Articles in peer-reviewed Journals

Auel, H., Ekau, W. (2009). Distribution and respiration of the high-latitude pelagic amphipod Themisto gaudichaudi in the Benguela Current in relation to upwelling. Progress in Oceanography, 83, 237-241.

Bode, M., Kreiner, A., van der Plas, A.K., Louw, D.C., Horaeb, R., Auel, H., Hagen, W. (2014). Spatio-temporal variability of copepod abundance along the 20°S monitoring transect in the northern Benguela Upwelling system from 2005 to 2011. PLoS ONE 9(5): e97738.

Bode, M., Schukat, A., Hagen, W., Auel, H. (2013). Predicting metabolic rates of calanoid copepods. Journal of Experimental Marine Biology and Ecology 444, 1-7.

Bohata, K., Koppelmann. R. (2013). Chaetognatha of the Namibian Upwelling Region: Taxonomy, Distribution and Trophic Position. PLoS ONE 8:e53839. doi:10.1371/journal.pone.0053839.

Burd, A., Koppelmann, R. et al. (2010). Assessing the apparent imbalance between geochemical and biochemical indicators of meso- and bathypelagic biological activity: What the @$#! is wrong with present calculations of carbon budgets? Deep-Sea Research II 57, 1557 - 1571.

Ekau, W., Auel, H., Hagen, W., Koppelmann, R., Wasmund, N., Bohata, K., Buchholz, F., Geist, S., Martin, B., Schukat, A., Verheye, H.M., Werner, T. (2018). Pelagic key species and mechanisms driving energy flows in the norther Benguela upwelling ecosystem and their feedback into biogeochemical cycles. Journal of Marine Systems 188, 49-62.

Ekau, W., Auel, H., Pörtner H.O., Gilbert D (2010). Impacts of hypoxia on the structure and processes in pelagic communities (zooplankton, macro-invertebrates and fish). Biogeosciences 7, 1669–1699.

Emeis, K., Eggert, A., Flohr, A., Lahajnar, N., Nausch, G., Neumann, A., Rixen, T., Schmidt, M., Van der Plas, A., Wasmund, N. (2018): Biogeochemical processes and turnover rates in the Northern Benguela Upwelling System. Journal of Marine Systems 188, 63-80.

Fennel, W., Junker, T., Schmidt, M., Mohrholz, V. (2012). Response of the Benguela Upwelling Systems to spatial variations in the wind stress. Continental Shelf Research 45, 65-77.

Flohr, A., van der Plas, A. K., Lahajnar, N., Neumann, A., Emeis, K.-C., Rixen, T. (submitted). Total alkalinity production buffers the carbonate saturation state in the Benguela Current upwelling system.

Flohr, A., van der Plas, A.K., Emeis, K.-C., Mohrholz, V., Rixen, T. (2014). Spatio-temporal patterns of C : N : P ratios in the northern Benguela upwelling regime. Biogeosciences 11, 885 - 897.

Geist, S.J., Verheye, H.M., Kunzmann, A., Ekau, W. (submitted). Larval feeding ecology of clupeoid fish in the degraded northern Benguela Current upwelling system and its implications for stock recruitment processes.

Geist, S. J., Kunzmann, A., Verheye, H. M., Eggert, A., Schukat, A., and Ekau, W. (2014). Distribution, feeding behaviour, and condition of Cape horse mackerel early life stages, Trachurus capensis, under different environmental conditions in the northern Benguela upwelling ecosystem. ICES Journal of Marine Science. doi:10.1093/icesjms/fsu087.

Geist, S.J., Kunzmann, A., Ekau, W. (2013). Energy demand of larval and juvenile Cape horse mackerels, Trachurus capensis, and indications for hypoxia tolerance as benefit in a changing environment. Marine Biology 160 (12), 3221-3232.

Grote, B., Stenevik, E.K., Ekau, W., Verheye, H.M., Lipiński M.R., Hagen, W. (2012). Spawning strategies and transport of early stages of the two Cape hake species, Merluccius paradoxus and M. capensis, in the southern Benguela upwelling system. African Journal of Marine Science, 34, 195-204.

Grote, B., Ekau, W., Stenevik E., Clemmesen, C., Verheye, H., Lipinski, M., Hagen, W. (2012). Characteristics of survivors: growth and nutritional condition of early stages of the hake species Merluccius paradoxus and M. capensis in the southern Benguela ecosystem. ICES Journal of Marine Science, 69, 553-562.

Grote, B., Hagen, W., Lipinski, M., Verheye, H., Stenevik, E., Ekau, W. (2011). Lipids and fatty acids as indicators of egg condition, larval feeding and maternal effects in Cape hakes (Merluccius paradoxus and M. capensis). Marine Biology 158, 1005-1017.

Gutknecht, E., Dadou, I., Le Vu, B., Cambon, G., Sudre, J., Garcon, V., Machu, E., Rixen, T., Kock, A., Flohr, A., Paulmier, A., Lavik, G. (2013). Coupled physical/biogeochemical modeling including O₂-dependent processes in the Eastern Boundary Upwelling Systems: Application in the Benguela. Biogeosciences 10, 3559-3591.

Gutknecht, E., Dadou, I., Le Vu, B., Marchesiello, P., Cambon, G., Sudre, J., Garcon, V., Machu, E., Rixen, T., Kock, A., Flohr, A., Paulmier, A., Lavik, G. (2011). Nitrogen transfers and sea-air N₂O fluxes in the upwelling off Namibia within the oxygen minimum zone: a 3-D model approach. Biogeosciences Discussion 8, 3537-3618. 

Hansen, A., Ohde, T., Wasmund, N. (2014). Succession of micro- and nanoplankton groups in ageing upwelled waters off Namibia, Journal of Marine Systems 140, 130-137.

Herbert, N.A., Goodman, M., Kunzmann, A. (2012). The low O₂ avoidance strategy of the cape silverside Atherina breviceps (Teleostei). Marine Freshwater Behaviour and Physiology 45 (3), 199-208.

Herzfeld, M., Schmidt, M., Griffies, S.M., Liang, Z. (2011). Realistic test cases for limited area ocean modeling. Ocean Modelling 37, 1–34.

Huenerlage, K., Buchholz, F. (2012). Krill of the northern Benguela Current and the Angola-Benguela frontal zone compared: physiological performance and short-term starvation in Euphausia hanseni. Journal of Plankton Research 35, 337-351.

Junker, T., Schmidt, M., Mohrholz, V. (2015). The relation of wind stress curl and meridional transport in the Benguela upwelling system. Journal of Marine Systems 143, 1–6.

Koppelmann, R., Jung, S.A., Lahajnar, N. (2014). Abundance and trophic position of gelatinous zooplankton in the Namibian upwelling region. In: Zooplankton: Species Diversity, Distribution and Seasonal Dynamics. Editor: George Kehayias. Nova Science Publishers, Inc. New York. pp 177-207.

Koppelmann, R., Kullmann, B., Lahajnar, N., Martin, B., Mohrholz, V. (2013). Onshore-offshore distribution of Thecosomata (Gastropoda) in the Benguela Current Upwelling Region off Namibia: species diversity and trophic position. Journal of the Marine Biological Association of the United Kingdom 93, 1625-1640.

Kreiner, A., Ekau, W., Stenevik, E. (2009). Sardine Sardinops sagax and anchovy Engraulis encrasicolus larvae avoid regions with low dissolved oxygen concentration in the northern Benguela Current system. Journal of Fish Biology 74, 270-277.

Levin, L., Ekau, W., Gooday, A., Middelburg, J., Naqvi, S.W.A., Rabalais, N., Zhang, J. (2009). Effects of natural and human-induced hypoxia on coastal benthos. Biogeosciences 6, 2063-2098.

Louw, D., van der Plas, A., Mohrholz, V., Wasmund, N., Junker, T., Eggert, A. (2016). Seasonal and interannual phytoplankton dynamics and forcing mechanisms in the Northern Benguela upwelling system. Journal of Marine Systems 157, 124-134.

Martin B., Koppelmann R., Kassatov P. (accepted 2016): Ecological relevance of salps and doliolids in the northern Benguela Upwelling System. Journal of Plankton Research., doi.org/10.1093/plankt/fbw095

Martin, B., Eggert, A., Koppelmann, R., Diekmann, R., Mohrholz, V., Schmidt, M., , (2014). Spatio-temporal variability of mesozooplankton biomass and environmental control in the northern Benguela Upwelling System: field investigations and model simulation. Marine Ecology 35, DOI: 10.111/maec.12173.

Meisel, S., Struck, U., Emeis, K.-C. (2011). Nutrient regime and upwelling in the northern Benguela since the middle Holocene in a global context - a multi-proxy approach. Fossil Record 14, 153-169.

Meisel, S., Struck, U., Emeis, K.-C., Kristen, I. (2011). Nutrient dynamics and oceanographic features in the central Namibian upwelling region as reflected in delta N-15-signals of suspended matter and surface sediments. Fossil Record 14, 171-193.

Mitra A., Jónasdóttir S.H., Flynn K.J., Castellani C., Gentleman W., Halsband C., Licandro P., Agersted M.D., Calbet A., Lindeque P., Koppelmann R., Møller E.F., St. John M. (2014). Bridging the gap between marine biogeochemical and fisheries sciences; configuring the zooplankton link. Progress in Oceanography, DOI: 10.1016/j.pocean.2014.04.025

Mohrholz, V., Eggert, A., Junker, T., Nausch, G., Ohde, T., Schmidt, M (2014). Cross shelf hydrographic and hydrochemical conditions and its short term variability at the northern Benguela during a normal upwelling season. Journal of Marine Systems, dx.doi.org/10.1016/j.jmarsys.2014.04.019.

Muller, A.A., Reason, Ch.J., Mohrholz, V., Schmidt, M., Eggert, A. (2014). Computing transport budgets along the shelf and across the shelf edge in the northern Benguela during summer (DJF) and winter (JJA). Journal of Marine Systems 140, 82-91.

Muller, A.A., Mohrholz, V., Schmidt, M. (2013). The circulation dynamics associated with a northern Benguela upwelling filament during October 2010. Continental Shelf Research 63, 59-68.

Nagel, B., Emeis, K.C., Flohr, A., Rixen, T., Schlarbaum, T., Mohrholz, V., v. d. Plas, A. (2013). N-cycling and balancing of the N-deficit generated in the oxygen minimum zone over the Namibian shelf - an isotope-based approach. Journal of Geophysical Research: Biogeosciences 118, doi: 10.1002/jgrg.20040.

Nagel, B., Gaye, B., Lahajnar, N., Struck, U., Emeis, K.-C. (2016). Effects of current regimes and oxygenation on particulate matter preservation on the Namibian shelf: Insights from amino acid biogeochemistry. Marine Chemistry 186, 121-132.

Neumann, A., Lahajnar, N., Emeis, K.-C. (2016). Benthic remineralisation rates in shelf and slope sediments or the northern Benguela upwelling margin. Continental Shelf Research 113, 47-61.

Ohde, Th., Mohrholz, V. (2011).  Interannual variability of Sulphur plumes off  the Namibian coast.  International Journal of Remote Sensing 32(24),  9327-9342.

Robinson, C., Koppelmann, R. et al. (2010). Mesopelagic zone ecology and biogeochemistry - a synthesis. Deep-Sea Research II 57, 1504-1518.

Schukat, A., Bode, M., Auel, H., Carballo, R., Martin, B., Koppelmann, R., Hagen, W. (2013). Pelagic decapods in the northern Benguela upwelling system: Distribution, ecophysiology and contribution to active carbon flux. Deep-Sea Research I 75, 146-156.

Schukat, A., Teuber, L., Hagen, W., Wasmund, N., Auel, H. (2013). Energetics and carbon budgets of dominant calanoid copepods in the northern Benguela upwelling system. Journal of Experimental Marine Biology and Ecology 442, 1-9.

Schukat, A., Auel, H., Teuber, L., Lahajnar, N., Hagen, W. (2014). Complex trophic interactions of calanoid copepods in the Benguela upwelling system. Journal of Sea Research 85, 186-196.

Teuber, L., Schukat, A., Hagen, W., Auel, H. (2014). Trophic interactions and life strategies of epi- to bathypelagic calanoid copepods in the tropical Atlantic Ocean. Journal of Plankton Research 36(4), 1109-1123.

Teuber, L., Schukat, A., Hagen, W., Auel, H. (2013). Distribution and ecophysiology of calanoid copepods in relation to the oxygen minimum zone in the Eastern Tropical Atlantic. PLoS ONE 8(11): e77590.

Tim, N., Zorita, E., Hünicke, B. (2015). Decadal variability and trends of the Benguela Upwelling System as simulated in a high-resolution ocean simulation. Ocean Science 11, 483-502. [pdf]

Tim, N., Zorita, E., Hünicke, B., Yi, X., Emeis, K.-C. (2016). The importance of external climate forcing for the variability and trends of coastal upwelling in past and future climate. Ocean Science 12, 807-823. [pdf]

Tremblay, N. , Werner, T. , Hünerlage, K. , Buchholz, F. , Abele, D. , Meyer, B., Brey, T. (2014). Euphausiid respiration model revamped: latitudinal and seasonal shaping effects on krill respiration rates. Ecological Modelling 291, 233-241, doi:10.1016/j.ecolmodel.2014.07.031.

Vorrath, M.-E., Lahajnar, N., Fischer, G., Libuku, V.M., Schmidt, M., Emeis, K.-C. (2018). Spatiotemporal variation of vertical particle fluxes and modelled chlorophyll a standing stocks in the Benguela Upwelling System. Journal of Marine Systems 180, 59-75.

Wasmund, N., Nausch, G., Hansen, A. (2014). Phytoplankton succession in an isolated upwelled Benguela water body in relation to different initial nutrient conditions, Journal of Marine Systems 140, 163-174.

Wasmund, N., Struck, U., Hansen, A., Flohr, A., Nausch, G., Grüttmüller, A., Voss, M. (2015). Missing nitrogen fixation in the Benguela region. Deep-Sea Research I 106, 30-41.

Wasmund, N., Siegel, H., Bohata, K., Flohr, A., Hansen, A., Mohrholz, V. (2016). Phytoplankton stimulation in frontal regions of Benguela upwelling filaments by internal factors. Frontiers in Marine Science, 3:210, doi: 10.3389/fmars.2016.00210.

Werner, T., Buchholz, C., Buchholz, F. (2015). Life in the sea of plenty: Seasonal and regional comparison of physiological performance of Euphausia hanseni in the northern Benguela upwelling system. Journal of Sea Research 103, 103-112.

Werner, T., Buchholz, F. (2013). Diel vertical migration behaviour in Euphausiids of the northern Benguela current: seasonal adaptations to food availability and strong gradients of temperature and oxygen. Journal of Plankton Research 35(4): 792-812.

Werner, T., Huenerlage, K., Verheye, H., Buchholz, F. (2012). Thermal constraints on the respiration and excretion rates of krill, Euphausia hanseni and Nematoscelis megalops, in the northern Benguela upwelling system off Namibia. African Journal of Marine Science 34, 391-399.

Wilhelm, M.R., Moloney, C., Paulus, S.C., Kashava, S., Brinkman, F.R.V., van der Plas, A.K., West, W.M., Jarre, A., Roux, J.-P. (2018): Biannual otolith zonation of Cape hake (Merluccius capensis) in response to fish physiology and environment in the northern Benguela. Journal of Marine Systems 188, 98-108.

Zhang, J., Gilbert, D., Gooday, A., Levin, L., Naqvi, S., Middelburg, J., Scranton, M., Ekau, W., Dewitte, B., Monteiro, P., Urban, E., Rabalais, N., Ittekkot, V., Kemp, W., van der Plas, A. (2010). Natural and human-induced hypoxia and consequences for coastal areas: synthesis and future development. Biogeosciences 7, 1443-1467.